The fundamental cause of the Yule-Nielsen effect, also called Optical Dot Gain, is the lateral scattering of light in paper. The lateral scatter of light can be described quantitatively by a point spread function, but a simpler model based on mean level scattering probabilities,
J.S. Arney, Cindy Scigaj, Prashant Mehta, "Linear Color Addition in Halftones" in Journal of Imaging Science and Technology, 2001, pp 466 - 471, https://doi.org/10.2352/J.ImagingSci.Technol.2001.45.5.art00008